Signa Farrow is the 19-year-old protagonist of Belladonna. Signa comes from a wealthy family and is set to inherit her family’s wealth and property when she comes of age, as both of her parents died when she was a baby. Signa lived with her loving grandmother for a few years before her natural death. After that, Signa was passed between less caring relatives who were only interested in the stipend that came with Signa’s guardianship and the hope that she would take care of them once she was given her inheritance. All of these relatives have untimely deaths, leading people in her community to believe that Signa is cursed or that she is a witch.

Having few people to instruct her or take her out into society, Signa is relatively sheltered and socially inexperienced compared to other girls her age, something she is very insecure about. She relies on the rules of etiquette and the limited expectations for women in society to guide her, not recognizing how superficial these ideas are until late in the novel. She dreams of marrying once she gets possession of her property at Foxglove Manor, something she is excited about as she believes it will prevent her from feeling as lonely as she has throughout her life.